The two tubes with globalfeedback around them forms one amplifier.
And you should look at them as one.
Their purpous is to produce gain with good distortion, flat freq response and low output impedance.
The outputtransformer is sensitive frequencies below its bandwidth produced by its inductance and plate impedance.
If an hp is to be inserted i recomend you to put it before the amp or perhaps after the OT... so that LF distortion is kept low.
Hope i got it all right.