Paul, what are you using for C4? If adjustable, try tightening it down all the way. This might kill any osc. that you have.
Everything looks halfway normal in the GR circuit, which is good news.
It looks like the trouble starts at the BH7 top plate. Before when you had 7 volts here, that would definately cause distortion because you are exeeding the 4 volt bias on the bottom cathode.
Try a 0.01 uF cap at 200 volts if you have one. Put it from ground to the bottom plate of V2 (pin 1) and see if that helps get that pin 6 ac volts reading down.
Also, what are you using for R16? Make sure it is 68K.
Do you have a Fender guitar amp around by any chance? If so, pull the 12AT7 phase inverter tube out and try it in the place of V1 and see what you get.
I am with AP here, since we have it narrowed down to the output stage, and your GR seems to be working good, I will do a new set of voltages with no GR. I will take home the signal generator to get a steady tone instead of music.
cj