You can run the unit with 3 tubes if you don't use the compressor. The other tubes have to be in place though for it pass signal.
From left side of chassis to right it's preamp tube, eq tube, output tube and compressor tube.
Also check the following 5600-7372 card. I think that is the correct card number. it is the one to the left of the PSU chassis. Check it for any failures. recap the 100UF/50V(blue vishay) and 10uF/63V(blue vishay)to extend life of your unit. I usually recap them with Panasonics even though they are not the same footprint, they work well. Come to think of it, that should be done on the eq and compressor cards too.