Freeze time with the top tube heater getting driven by the positive peak of the 6.3 volt supply.
If the heaters are wired correctly, the bottom tube heater will be driven by the same positive peak.
Any hum that drifts from the heater to the grid and gets amplified by the tubes will cancel out inside the output transformer.
This is because you will have a positive peak on the top of transformer primary and a positive peak on the bottom winding. Since there is no potential difference between the top and bottom of the primary, no heater him will be induced into the secondary and it's speaker.
This is confusing because of the top and bottom peaks getting cancelled by twisting the wire in the heater wiring, but in the output transformer, you want a difference in voltage across the primary if you are trying to amplify a guitar, as a potential difference across the primary will mean that current will flow.
Since the transformer is a current operated device, with no voltage difference between the two terminals of the winding as in our correctly wired heaters, no current will flow and so no hum is heard.
This is also why you do not need a ton of filtering in a push pull amp for the power tubes , the transformer will do the same thing with the power supply ripple as it does with the heater hum voltage.