The Q1 numbers are good. With your cathode resistor numbers, you take the voltage across each.
The -1.85V at pin 3 is the RV3 trimmed voltage less 1 diode drop of 0.6V at the Q1 base-emiiter junction.
eg. r1 -0.88vdc at one end, pin3[-1.85vdc] at the other, so current is 0.97V/330R -> 2.94mA
Similar for the others 2.36mA, 7.94mA, 3.12mA to give you the total +135V rail current draw.
Your R3 value of +0.77Vdc is very different from the others - I would say check that again.
Assuming that R1, R2, R4 areOK and R3 is the odd man out, that would imply a total current approx 6mA per signal phase phase and 12mA for the signal amp. (not sure if you have 1 signal amp pcb or 2 in this build).
It's a bit lowish but never mind for the moment - you should surely see some output at T2 secondary with 12mA flowing thru it.
The other thing to note is that the GR tubes conduct current according to the voltage difference between the grid and cathode. As mentioned before, in the case of no-signal idle, grid is held at a -ve reference voltage set by RV6 via the CV feed to T1 secondary. That should be around -4.5V as previosuly discussed.
So the bias applied to each of the GR tubes is grid -4.5V minus the cathode voltage -> -3.62V, -3.43V, -5.27V, -3.68V respectively. Cathode 3 is the odd man out (if your measurement is correct).
Looking at the data sheets for 6BC8 with plate of 135V and around -3.5V bias says around 3mA per stage is expected.
So you are pretty much as expected.
Check again that funny R3 number. If it is correct, you may have a dud tube section.
Move it around and see if the funny cathode voltage follows it.
If so, replace it, if not - check the resistor is actually 330R and not some other value and finally check solder for any issues.
They check again.
The reason I said a higher total signal amp current is that I bias mine to idle quite a bit higher - like 6mA (-2.5B bias) .
That's to give more available GR (more current = more gain therefor more room to compress before crapping out due to low current under GR) , but at the expense of input signal capability at idle (will clip the input at idle at lower inputs - bad) as well as higher idle current dissipation, which can be an issue with heat on tubes, regulators etc.
Not to mention higher noise when not compressing much. AND that the output traffo can handle that amount of dc CT fed thru it
To summarise, that looks more or less OK except for the bad R3 reading. Fix that up them satisfy yourself you get some good signal at xlr output connector then you can start looking at GR issues with the control amp.
Simple
Cheers
Alex