> effects of tying the two screens (G2) together fed with 100k
Well, you know the effect. Vari-Mu happens by reducing cathode current in T1 T2. Cathode current splits into plate and screen current, and this ratio is fairly fixed. Screens sit at 40V from a 170V rail, so there is large drop in the 100K. Screen current falls, drop is less, screen voltage rises.
So as gain goes down, screen voltage goes up. Why?
The maximum undistorted input on a pentode is roughly Vs/Mu. Speaking over-simple, as we get big signals, screen voltage rises to handle them. It is a very real effect. You find a similar condition on many remote-cutoff tubes for IF and RF service.
Offhand, E99F Mug2g1 is 27, so we expect 40V/27= 1.5V to just-distort at idle, say 150V/27= 5V to just-distort in deep GR.
This is also true on triodes, and many classic triode vari-Mus have large plate resistors and sit at low idle plate voltage.
It is not that simple, and you can also get good results with fairly fixed plate or screen voltage.
> "ac" - cathode bias of the vari mu pentodes
The vari-Mu stage has the lowest signal, as low as a mike-amp (I mean a classic mike-amp, not a rock-studio input). Here we need two tubes with low heater-hum, low hiss, and good matching. By force-nulling the hum with +/-8mVAC hum from pot 66, we may relax the requirement for low heater-hum, and just try to find two matched low-hiss tubes.
It isn't exactly the same as gitar amps. The usual plan is you twist your heater wires 99% symmetrical, so their radiated hum field is canceled at nearby plate/grid parts, and then tune your "center" tap a few percent off-center to match your few-percent asymmetric twisting. The limiter is a simpler case.
There is more to the tale. Note that T1 T2 are fed one-side-grounded 6.2VAC. In most low-level work, that would be wrong. I bet these Germans laid-out the T1 T2 heater wiring so exACTly that hum in the two tubes cancels 99% perfect. Then pot 66 trims that last dab.
Pot 61 adjusts thump. It will affect hum, but that's not what you should adjust it for. You probably have to go back and forth from 66 to 61 to 66 a few times to get both balances good.