I know a little bit about Helios, from reading and my experience building bits and pieces.
As far as I can tell (and putting aside the germanium discussion for the moment), the earlier Helios consoles (with the M10 input xfm) used the three transistor design amp (and slight variants thereof) as mic pre, eq make up, fader buffer, bus/master amp and for line outs. The amp was also found in fx returns. No two consoles were quite the same. The early consoles had low headroom (0 - 24V) and I believe they were preferably run as close to the edge of headroom as possible. There was probably quite a lot of subtle clipping going on (multiple stages - clipped signals being clipped again?) with possibly slightly starved outputs (e.g. the load on the buffer amp with the panpot, aux sends, &c). I read somewhere that the router contributed quite a bit to the overall console functioning/sound.
Later, the seven transistor design amp was used (32V - 36V) as the eq make up and fader buffer amp, which improved headroom but there was a change of input transformer that was not appreciated by all.
Given the number of amp stages, and the summing, it would be expected for such a console to have a different sound to just the mic pre eq? Perhaps even a channel strip (with router) has a different sound to a mic pre eq?
It is said (in the wisdom of the interwebs) that early Helios consoles were noisy, high cross talk, and had low headroom and that it was all down to the talent that was recorded (who would have sounded good recorded with a 4-track Tascam porta studio).
Personally, I happen to like the sound and the simplicity of the Helios designs.