It is not significantly bootstrapped. The transformer secondary is AC-grounded at the 0.25u cap.
DC bias may be adjusted without significant effect on the AC gain.
V1a bias is just the 120 ohm resistor. This gets to its grid through a 160K/0.25uFd filter, to remove signal and to hold the "cold" end of the transformer winding at AC ground. It is not directly affected by voltage across the 1500 ohm resistor. It is indirectly affected because V across 1500 subtracts from V1a plate-cathode voltage, but for any reasonable values it causes no great change in V1a performance.
The AC gain is roughly (17.5K/(120+1500))+1 or just over 11. The 1500 ohm may be adjusted over a wide range.
It can be set to zero to maximize AC gain. This will increase V1a current about 5%, and V2 current about 8%, neither is a problem. AC gain would be near 145. This is still a High-NFB amp with useful input overload. If you want "more", shunt the 120 ohm with a big electrolytic. Assuming 1:10 iron, input overload is 3mV.
The 1500 could be made larger, though for large increase you might want to reduce the 17.5K in addition or instead, to keep V2 current similar. If you changed 17.7K:1500 to 10K:10K, the AC gain would be 2, and V1a current would be down 25%. Trying to get closed-loop gain down to unity would require re-thinking... but at that point it would be whole-lot-simpler to make V1 and V2 both 12AU7 as quad cathode-follower.
The top of the 0.25u sits at about +14V as drawn, at zero with the 1500 shorted, and near +80V with a 10K:10K NFB loop. Compared with voltage breakdown limits on the 0.25u, the transformer, and V1a heater, this is safe.
The dead men knew what they were doing. Often including the idea that gain might have to be changed per customer needs, without re-design effort.