I was referring to the design of the original post, not the DESA one.
Having a 16.5K cathode resistor, not bypassed, is going to raise the total resistance of the output tubes, in relation to the transformer.
The 16.5K resistor will compensate this through negative feedback, which is a form of correction (that is what I meant), and the amount of correction needed because of the high resistance driving the reactive load, is probably more than it should be, without affecting the sound, even though THD may be low enough.
Also, such a high value for a common resistor between cathodes, appears to be doing good things for reducing distortion in a push-pull class A circuit, but since the tubes used are usually not matched perfectly, and don't have perfectly equal curves, the phase relationship between the tubes, will also be mismatched to some degree, and instead of cancelling out harmonics, you get positive feedback of odd harmonics, between the tubes.
There's a paper about this, relative to power tube output, but I think that to some degree it applies to push-pull tube class-A in general, especially when the tubes are loaded heavily.
The argument being that there should be no interaction between the two output tubes, via the cathode resistor, because of the cross-modulation between the tubes, and that it generally increases Intermodulation Distortion, while it can lower THD. So the idea is to bypass the cathode resistor and just use more local feedback on either side, to reduce distortion.