Ben,
The input amplifier is DC decoupled by C1, C5 and C9. Somewhere in this 3-transistor amplifier is a wrong component or faulty transistor. It's about a dozen resistors and 3 transistors. Did you replace Q4 at all? Short of having the unit if front of me to examine and probe, I can only suggest the obvious areas to check. You will need to re-check all of these components carefully. Something may be causing the whole section to oscillate (which you will not see on your multimeter) so double-check the capacitors (C2, C6, C5, C8) just in case.
BTW, I have a circuit that I drew of my unit, which is much the same as Mnats Rev J except I added a couple more compression ratios and did not use the on-board PSU. It may help you during fault finding.
http://www.axtsystems.com/index.php?view=article&catid=34%3A1176ln&id=45%3A1176lncircuits&option=com_content&Itemid=62
Geoff